But an inclusive fitness perspective suggests this is wrong-headed because they were never different individuals in the first place – they were always of one genotype, and the multicellular organism is just a phenotype of this singular evolutionary unit. Problem solved?

Entities in the same semantic neighborhood but without a typed relation to this one — candidates for new edges or unrecognized duplicates.

• For a new level of individuality to have causal role as evolutionary unit, heritable fitness differences must be non-aggregative (not decomposable into lower-level differences).claim0.793
  Core claim: monotonic non-linearities are insufficient; evolutionary outcomes must change depending on context.
• Target morphology shifts occur despite the fact that all of the individual cells have unaltered normal genomes, showing that competent subunits can be pushed to implement diverse organism-scale goals by physiological signals.claim0.787
  Highlights the non-genetic control of large-scale anatomy.
• The kind of relationships necessary to produce evolutionary individuality – the generation and heritability of fitness differences at the collective level – are the same as those required to produce organismic individuality – the information integration and collective action characteristic of a self.claim0.786
  Core conjecture linking evolutionary and organismic individuality.
• Organismic individuality can be separated from genetics: integration and collective action occur in non-neural systems.finding0.785
  Empirical findings from developmental biology (Manicka & Levin, Lyon et al.) supporting mechanistic basis for individuality independent of genetic determination.
• For collectives to be meaningful evolutionary units, fitness interactions must be non-linearly separable, not merely non-aggregative.claim0.779
  Strengthens distinction between monotonic non-linear interactions (explanatorily redundant) and non-decomposable interactions (causally significant).
• Multi-scale competency likely explains the remarkable evolvability of living forms because it flattens the fitness landscape, allowing mutations' negative effects to be buffered while positive effects accumulate.claim0.775
  Mechanistic claim connecting multi-scale competency architecture to evolutionary robustness
• Genetic homogeneity is neither necessary nor sufficient for organismic individuality; functional integration is more importantclaim0.770
  Interpretive claim supported by experimental findings on planaria, chimeras, and cancer
• Biology's robustness, open-endedness, evolvability, and unique complexity likely depend on the fact that evolution works with an agential material.claim0.769
  Central claim linking life's properties to the inherent competencies of its material substrate.